Stephen Richards, Ph.D.

image: Stephen Richards, Ph.D.Contact Information

srichards - at -

Dr. Richards has now moved to UC Davis. Please contact him with any questions about the arthropod projects.


Ph.D., Rice University, 1999
B.Sc., The University of Bath, 1990

Research Interests:

Dr. Stephen Richards has a broad background in genomics, with a focus on quantitative traits in insect models, arthropod comparative genomics, methods for genome sequencing, assembly and annotation, and a history in human genetics.


Fuller ZL, Haynes GD, Zhu D, et al. Evidence for stabilizing selection on codon usage in chromosomal rearrangements of Drosophila pseudoobscura. G3 (Bethesda). 2014;4(12):2433-49. doi:10.1534/g3.114.014860.

Chen Z-X, Sturgill D, Qu J, et al. Comparative validation of the D. melanogaster modENCODE transcriptome annotation. Genome Res. 2014;24(7):1209-23. doi:10.1101/gr.159384.113.

Huang W, Massouras A, Inoue Y, et al. Natural variation in genome architecture among 205 Drosophila melanogaster Genetic Reference Panel lines. Genome Res. 2014;24(7):1193-208. doi:10.1101/gr.171546.113.

Chipman AD, Ferrier DEK, Brena C, et al. The first myriapod genome sequence reveals conservative arthropod gene content and genome organisation in the centipede Strigamia maritima. PLoS Biol. 2014;12(11):e1002005. doi:10.1371/journal.pbio.1002005.

Bradnam KR, Fass JN, Alexandrov A, et al. Assemblathon 2: evaluating de novo methods of genome assembly in three vertebrate species. Gigascience. 2013;2(1):10. doi:10.1186/2047-217X-2-10.

Ober U, Ayroles JF, Stone EA, et al. Using whole-genome sequence data to predict quantitative trait phenotypes in Drosophila melanogaster. PLoS Genet. 2012;8(5):e1002685. doi:10.1371/journal.pgen.1002685.

Fondon JW, Martin A, Richards S, Gibbs RA, Mittelman D. Analysis of microsatellite variation in Drosophila melanogaster with population-scale genome sequencing. PLoS One. 2012;7(3):e33036. doi:10.1371/journal.pone.0033036.

Mackay TFC, Richards S, Stone EA, et al. The Drosophila melanogaster Genetic Reference Panel. Nature. 2012;482(7384):173-8. doi:10.1038/nature10811.

English AC, Richards S, Han Y, et al. Mind the gap: upgrading genomes with Pacific Biosciences RS long-read sequencing technology. PLoS One. 2012;7(11):e47768. doi:10.1371/journal.pone.0047768.

Huang W, Richards S, Carbone MAnna, et al. Epistasis dominates the genetic architecture of Drosophila quantitative traits. Proc Natl Acad Sci U S A. 2012;109(39):15553-9. doi:10.1073/pnas.1213423109.